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In this anti- romantic comedy about love and fate, a young greeting card writer (Joseph Gordon-Levitt) is hopelessly, helplessly searching for the girl of his dreams...and his new co-worker, Summer Finn (Zooey Deschanel), may just be "the one." But the 500 days of their offbeat relationship reveal that the road to happiness can be unpredictable, uncontrollable and unbelievable.